Figure P58

5.9. This problem will model the three-ommatidium LI system described by Equation 5.3-2 and Figure 5.3-5. A block diagram for this system is shown in Figure P5.9. Note that LI is modeled here by a feed-forward architecture. The retinula cells of each ommatidium respond to light by depolarizing, but not spiking. A single eccentric cell associated with each ommatidium produces spikes as a result of depolarization excitation being coupled to it from its surrounding retinula cells (see Figure 5.3-1).

f(x) = (I„/2)|l - sin(uox)l f(x) = (I„/2)|l - sin(uox)l s(x) = exp( a' x2 )

Retinula cell 1

Retinula cell 2

Retinula cell 3

—' K,3 K12 —• K21 ^y1 K23 l— — K32 K31 —


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